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		Professor Naoki Koyama of Kyoto University began a major long-term study of individual ringtailed lemurs in 1989. This has resulted in articles on demography, female-female relations, troop fission and formation, and on the relation of lemur home ranges to the presence and persistence of tamarind trees. His students Shinichiro Ichino and Takayo Soma carry on the study to the present day, with many collaborators and students from Japan. Prof. Koyama has summarized much of his work and knowledge of Madagascar in a general book, (2009). Madagascar tou–Nishiindoyou chiikikennkyuu nyuumon (The Island of Madagascar: an introduction to Area Studies in the Western Indian Ocean Region). Kanagawa, Tokaidaigaku Press (in Japanese). Contact: "N.Koyama" mv2z67@bma.biglobe.ne.jp
Portrait of Professor Naoki Koyama © photo by A. Jolly

Professor Naoki Koyama of Kyoto University began a major long-term study of individual ringtailed lemurs in 1989. This has resulted in articles on demography, female-female relations, troop fission and formation, and on the relation of lemur home ranges to the presence and persistence of tamarind trees. His students Shinichiro Ichino and Takayo Soma carry on the study to the present day, with many collaborators and students from Japan. Prof. Koyama has summarized much of his work and knowledge of Madagascar in a general book, (2009). Madagascar tou–Nishiindoyou chiikikennkyuu nyuumon (The Island of Madagascar: an introduction to Area Studies in the Western Indian Ocean Region). Kanagawa, Tokaidaigaku Press (in Japanese).

		Shinichiro Ichino discussing field research with colleagues. © photo by A. Jolly

Shinichiro Ichino has studied ringtailed lemurs at Berenty since 2000, focusing on male mating, migration, and reproductive success. Contact: ichino@pri.kyoto-u.ac.jp Takayo Soma did her PhD at Berenty on the feeding behavior of a troop ringtailed lemurs which depended heavily on introduced plants. Her meticulous studies allowed calibration of the actual intake of nutrients from different sources over several years, as well as following the detailed demography of seven different troops, in collaboration with Shinichiro Ichino. Her supervisor was Naoki Koyama. Contact: soma@jambo.africa.kyoto-u.ac.jp
					Takayo Soma with a maki baby

Shinichiro Ichino can be reached at: Genome Diversity Section, Dept. of Evolution and Phylogeny Primate Research Institute Kyoto University 41-2 Kanrin Inuyama, Aichi, 484-8506 JAPAN Fax: +81-568-63-0565					Takayo Soma can be reached at: Center for African Area Studies Kyoto University 46 Yoshida Shimoadachi-cho Sakyo, Kyoto 606-8501, JAPAN Tel: +81-75-753-7800 Fax: +81-75-753-7810

Shinichiro Ichino has studied ringtailed lemurs at Berenty since 2000, focusing on male mating, migration, and reproductive success.